Imprinted Gene Databases

Imprinted Genes: by Species

Below are listings of genes by species, sorted by chromosomal location. Gene information has been gathered from NCBI, and some genes lack chromosomal coordinates; these are designated with ---. To switch species, select the appropriate tab. To view more information about a gene, click on its name. If there is a gene missing that you feel should be included in the catalog, please contact us.

Only display genes with imprint status:
Gene Aliases Location Status Expressed Allele
NDUFA4P1 NDUFA4  1p13.3  Predicted Paternal
GFI1 GFI-1, ZNF163, FLJ94509  1p22 AS  Predicted Paternal
DIRAS3 ARHI, NOEY2  1p31 AS  Imprinted Paternal
FUCA1 FUCA  1p34 AS  Predicted Paternal
RNU5D-1 U5DL, U5DS, RNU5D  1p34.1 AS  Imprinted Paternal
BMP8B OP2, BMP8, MGC131757  1p35-p32 AS  Predicted Paternal
DVL1 DVL, MGC54245  1p36 AS  Predicted Maternal
WDR8 FLJ20430, MGC99569  1p36.3 AS  Predicted Maternal
TP73 P73  1p36.3  Imprinted Maternal
RPL22 EAP, HBP15, HBP15/L22  1p36.3-p36.2 AS  Predicted Paternal
PRDM16 MEL1, PFM13, KIAA1675, MGC166915  1p36.23-p33  Predicted Paternal
PEX10 NALD, RNF69, MGC1998  1p36.32 AS  Predicted Maternal
TMEM52   1p36.33 AS  Predicted Paternal
HSPA6   1q23  Predicted Maternal
PTPN14 PEZ, PTP36, MGC126803  1q32.2 AS  Predicted Maternal
HIST3H2BB   1q42.13  Predicted Maternal
OBSCN UNC89, FLJ14124, KIAA1556, KIAA1639, MGC120409, MGC120410, MGC120411, MGC120412, MGC138590, DKFZp666E245  1q42.13  Predicted Paternal
OR11L1   1q44 AS  Predicted Paternal
LRRTM1   2p12 AS  Imprinted Paternal
VAX2 DRES93  2p13  Predicted Maternal
OTX1 FLJ38361, MGC15736  2p13  Predicted Maternal
COMMD1 MURR1, C2orf5, MGC27155  2p15  Not Imprinted Biallelic
CCDC85A KIAA1912  2p16.1  Predicted Paternal
ABCG8 GBD4, STSL, MGC142217  2p21  Predicted Maternal
CYP1B1 CP1B, GLC3A, P4501B1  2p21 AS  Predicted Paternal
ZFP36L2 BRF2, ERF2, ERF-2, TIS11D, RNF162C  2p22.3-p21 AS  Predicted Maternal
GPR1   2q33.3 AS  Imprinted Paternal
ZDBF2   2q33.3  Imprinted Paternal
TIGD1 EEYORE  2q37.1 AS  Predicted Paternal
MYEOV2   2q37.3 AS  Predicted Paternal
ALDH1L1 FTHFD, DKFZp781N0997  3q21.3 AS  Predicted Maternal
ZIC1 ZIC, ZNF201  3q24  Predicted Maternal
HES1 HHL, HRY, HES-1, bHLHb39, FLJ20408  3q28-q29  Predicted Paternal
FGFRL1 FHFR, FGFR5  4p16  Predicted Maternal
KIAA1530 MGC117169  4p16.3  Predicted Maternal
SPON2 DIL1, DIL-1, Mindin, M-spondin  4p16.3 AS  Predicted Paternal
NAP1L5 DRLM  4q22.1 AS  Imprinted Paternal
GAB1   4q31.21  Not Imprinted Biallelic
ADAMTS16 FLJ16731, ADAMTS16s  5p15  Predicted Maternal
CDH18 CDH14, CDH24, CDH14L, EY-CADHERIN  5p15.2-p15.1 AS  Predicted Paternal
CSF2 GMCSF, MGC131935, MGC138897  5q31.1  Predicted Maternal
PRIM2 p58, PRIM2A  6p12-p11.1 AS  Conflicting Data Biallelic
BTNL2 SS2, BTL-II, HSBLMHC1  6p21.3 AS  Predicted Maternal
FAM50B X5L, D6S2654E  6p25.2  Imprinted Paternal
MRAP2 C6orf117, bA51G5.2, RP11-51G5.2  6q14.2  Predicted Paternal
C6orf117 RP11-51G5.2  6q14.3  Predicted Paternal
AIM1 ST4, CRYBG1  6q21  Imprinted Paternal
LIN28B CSDD2  6q21  Imprinted Paternal
PLAGL1 ZAC, LOT1, ZAC1, MGC126275, MGC126276, DKFZp781P1017  6q24-q25 AS  Imprinted Paternal
HYMAI NCRNA00020  6q24.2 AS  Imprinted Paternal
SLC22A2* OCT2, MGC32628  6q26 AS  Imprinted Maternal
IGF2R MPR1, MPRI, CD222, CIMPR, M6P-R  6q26  Not Imprinted Biallelic
SLC22A3* EMT, EMTH, OCT3  6q26-q27  Imprinted Maternal
BRP44L CGI-129, dJ68L15.3  6q27 AS  Predicted Paternal
GRB10 RSS, IRBP, MEG1, GRB-IR, Grb-10, KIAA0207  7p12-p11.2 AS  Imprinted Isoform Dependent
DDC AADC  7p12.2 AS  Imprinted Isoform Dependent
GLI3 PHS, ACLS, GCPS, PAPA, PAPB, PAP-A, PAPA1, PPDIV  7p13 AS  Predicted Maternal
HOXA5 HOX1, HOX1C, HOX1.3, MGC9376  7p15-p14 AS  Predicted Maternal
HOXA4 HOX1, HOX1D  7p15-p14 AS  Predicted Maternal
HOXA2 HOX1K  7p15-p14 AS  Predicted Maternal
HOXA11 HOX1, HOX1I  7p15-p14 AS  Predicted Maternal
EVX1   7p15-p14  Predicted Paternal
HOXA3 HOX1, HOX1E, MGC10155  7p15-p14 AS  Predicted Maternal
TMEM60 DC32, C7orf35, MGC74482  7q11.23 AS  Predicted Paternal
MAGI2 AIP1, AIP-1, ARIP1, SSCAM, MAGI-2, ACVRIP1  7q21 AS  Imprinted Maternal
PEG10 EDR, HB-1, Mar2, MEF3L, Mart2, RGAG3  7q21  Imprinted Paternal
SGCE ESG, DYT11  7q21-q22 AS  Imprinted Paternal
ASB4 ASB-4, MGC142039, MGC142041  7q21-q22  Not Imprinted Biallelic
PON2   7q21.3 AS  Not Imprinted Biallelic
PON3   7q21.3 AS  Not Imprinted Biallelic
CALCR CRT, CTR, CTR1  7q21.3 AS  Provisional Data Maternal
PON1 ESA, PON, MVCD5  7q21.3 AS  Provisional Data Maternal
PPP1R9A NRB1, NRBI, FLJ20068, KIAA1222, Neurabin-I  7q21.3  Imprinted Maternal
DLX5   7q22 AS  Imprinted Maternal
TFPI2 PP5, REF1, TFPI-2, FLJ21164  7q22 AS  Imprinted Maternal
COPG2 2-COP, FLJ11781, DKFZp761N09121  7q32 AS  Conflicting Data Paternal
COPG2IT1 CIT1, COPG2AS, FLJ41646, NCRNA00170, DKFZP761N09121  7q32  Imprinted Paternal
CPA4 CPA3  7q32  Imprinted Maternal
MEST PEG1, MGC8703, MGC111102, DKFZp686L18234  7q32  Imprinted Paternal
MESTIT1 MEST-IT, PEG1-AS, MEST-AS1, MEST-IT1, NCRNA00040  7q32.2 AS  Imprinted Paternal
KLF14 BTEB5  7q32.3 AS  Imprinted Maternal
FASTK FAST  7q35 AS  Predicted Maternal
SLC4A2 AE2, HKB3, BND3L, NBND3, EPB3L1  7q35-q36  Predicted Maternal
PURG PURG-A, PURG-B, MGC119274  8p11 AS  Predicted Paternal
DLGAP2 DAP2, SAPAP2  8p23  Imprinted Paternal
NKAIN3 FAM77D, FLJ39630  8q12.3  Predicted Paternal
LY6D E48  8q24-qter AS  Predicted Paternal
GPT AAT1, ALT1, GPT1  8q24.3  Predicted Maternal
KCNK9 KT3.2, TASK3, K2p9.1, TASK-3, MGC138268, MGC138270  8q24.3 AS  Imprinted Maternal
ZFAT-AS1 ZFATAS, ZFAT-AS, SAS-ZFAT, NCRNA00070  8q24.22  Imprinted Paternal
ZFAT AITD3, ZFAT1, ZNF406  8q24.22 AS  Imprinted Paternal
GLIS3 ZNF515  9p24.2 AS  Imprinted Paternal
APBA1 X11, X11A, MINT1, D9S411E, X11ALPHA  9q13-q21.1 AS  Predicted Paternal
C9orf85 MGC61599, RP11-346E17.2  9q21.13  Predicted Paternal
FLJ46321 FLJ46321  9q21.32  Predicted Maternal
LMX1B NPS1, LMX1.2, MGC138325, MGC142051  9q34  Predicted Maternal
PHPT1 PHP14, CGI-202, HSPC141, bA216L13.10, DKFZp564M173, RP11-216L13.10  9q34.3  Predicted Maternal
EGFL7 ZNEU1, MGC111117, VE-STATIN, RP11-251M1.2  9q34.3  Predicted Paternal
C9orf116 FLJ13945, MGC29761, RP11-426A6.4  9q34.3 AS  Predicted Paternal
SFMBT2   10p14 AS  Not Imprinted Biallelic
GATA3 HDR, MGC2346, MGC5199, MGC5445  10p15  Predicted Paternal
CTNNA3 VR22, MGC26194, MGC75041  10q22.2 AS  Provisional Data Maternal
LDB1 NLI, CLIM2  10q24-q25 AS  Predicted Maternal
NKX6-2 GTX, NKX6B, NKX6.2, MGC126684  10q26 AS  Predicted Maternal
VENTX NA88A, HPX42B, VENTX2, MGC119910, MGC119911  10q26.3  Predicted Maternal
C10orf93 bB137A17.3, RP13-137A17.3  10q26.3 AS  Predicted Maternal
C10orf91 bA432J24.4, RP11-432J24.4  10q26.3  Predicted Maternal
PAOX PAO, MGC45464, DKFZp434J245  10q26.3  Predicted Maternal
INPP5F V2 SAC2, hSAC2, MSTP007, MSTPO47, FLJ13081, KIAA0966, MGC59773, MGC131851  10q26.11  Imprinted Paternal
WT1-Alt trans WT1, GUD, WAGR, WT33, WIT-2  11p13 AS  Imprinted Paternal
PKP3   11p15  Predicted Maternal
KCNQ1OT1 LIT1, KvDMR1, KCNQ10T1, KvLQT1-AS, long QT intronic transcript 1  11p15  Imprinted Paternal
AMPD3   11p15  Not Imprinted Biallelic
OSBPL5 ORP5, OBPH1, FLJ42929  11p15.4 AS  Imprinted Maternal
KCNQ1DN BWRT, HSA404617  11p15.4  Imprinted Maternal
ZNF215 BAZ2, ZKSCAN11  11p15.4  Provisional Data Maternal
TSPAN32 ART1, PHMX, PHEMX, TSSC6, FLJ17158, FLJ97586, MGC22455  11p15.5  Not Imprinted Biallelic
CDKN1C BWS, WBS, p57, BWCR, KIP2  11p15.5 AS  Imprinted Maternal
TSSC4   11p15.5  Not Imprinted Biallelic
INS ILPR, IRDN  11p15.5 AS  Imprinted Paternal
B4GALNT4 FLJ25045  11p15.5  Predicted Maternal
PHLDA2 IPL, BRW1C, BWR1C, HLDA2, TSSC3  11p15.5 AS  Imprinted Maternal
TH TYH, DYT14, DYT5b  11p15.5 AS  Unknown Unknown
NAP1L4 NAP2, NAP2L, hNAP2, MGC4565, NAP1L4b  11p15.5 AS  Unknown Unknown
IFITM1 9-27, CD225, IFI17, LEU13  11p15.5  Predicted Maternal
KCNQ1 LQT, RWS, WRS, LQT1, SQT2, ATFB1, ATFB3, JLNS1, KCNA8, KCNA9, Kv1.9, Kv7.1, KVLQT1, FLJ26167  11p15.5  Imprinted Maternal
IGF2 INSIGF, pp9974, C11orf43, FLJ22066, FLJ44734  11p15.5 AS  Imprinted Paternal
TRPM5 MTR1, LTRPC5  11p15.5 AS  Provisional Data Paternal
CD81 S5.7, TAPA1, TSPAN28  11p15.5  Not Imprinted Biallelic
ASCL2 ASH2, HASH2, MASH2, bHLHa45  11p15.5 AS  Not Imprinted Biallelic
SLC22A18AS BWR1B, BWSCR1B, ORCTL2S, SLC22A1LS, p27-BWR1B  11p15.5 AS  Provisional Data Maternal
H19 ASM, BWS, ASM1, MGC4485, PRO2605, D11S813E  11p15.5 AS  Imprinted Maternal
IGF2AS PEG8, MGC168198  11p15.5  Imprinted Paternal
SLC22A18 HET, ITM, BWR1A, IMPT1, TSSC5, ORCTL2, BWSCR1A, SLC22A1L, p45-BWR1A, DKFZp667A184  11p15.5  Imprinted Maternal
RAB1B   11q12  Predicted Maternal
ANO1 DOG1, TAOS2, ORAOV2, TMEM16A  11q13.3  Imprinted Maternal
DHCR7 SLOS  11q13.4 AS  Not Imprinted Biallelic
ZC3H12C MCPIP3  11q22.3  Imprinted Paternal
KBTBD3 BKLHD3, FLJ30685  11q22.3 AS  Predicted Paternal
SDHD PGL, CBT1, PGL1, SDH4  11q23  Conflicting Data Paternal
NTM HNT, NTRI, IGLON2  11q25  Imprinted Maternal
ABCC9 SUR2, ABC37, CMD1O, FLJ36852  12p12.1 AS  Predicted Maternal
RBP5 CRBP3, CRBPIII, CRBP-III  12p13.31 AS  Imprinted Maternal
SLC26A10   12q13  Predicted Maternal
SLC38A4 ATA3, NAT3, PAAT, FLJ10191, MGC126876  12q13 AS  Unknown Unknown
HOXC9 HOX3, HOX3B  12q13.3  Predicted Maternal
HOXC4 HOX3, cp19, HOX3E  12q13.3  Predicted Maternal
CDK4 CMM3, PSK-J3, MGC14458  12q14 AS  Predicted Maternal
E2F7 FLJ12981  12q21.2 AS  Predicted Maternal
DCN CSCD, PG40, PGII, PGS2, DSPG2, SLRR1B  12q21.33 AS  Unknown Unknown
KIAA1545 KIAA1545  12q24.33  Predicted Maternal
FBRSL1 KIAA1545  12q24.33  Predicted Maternal
HTR2A HTR2, 5-HT2A  13q14-q21 AS  Conflicting Data Maternal
RB1 RB, pRb, OSRC, pp110, p105-Rb  13q14.2  Imprinted Maternal
FLJ40296 FLJ40296, MGC149404, MGC149405  13q21.1  Predicted Maternal
FAM70B MGC20579, RP11-199F6.1  13q34  Predicted Maternal
FOXG1 BF1, BF2, QIN, FKH2, HBF2, HFK1, HFK2, HFK3, KHL2, FHKL3, FKHL1, FKHL2, FKHL3, FKHL4, HBF-1, HBF-2, HBF-3, FOXG1A, FOXG1B, FOXG1C, HBF-G2  14q13  Predicted Paternal
FERMT2 MIG2, KIND2, mig-2, UNC112, PLEKHC1, UNC112B, FLJ34213, FLJ44462, DKFZp686G11125  14q22.1 AS  Predicted Paternal
DLK1 DLK, FA1, ZOG, pG2, PREF1, Pref-1  14q32  Imprinted Paternal
DIO3 D3, 5DIII, TXDI3, DIOIII  14q32  Unknown Unknown
MEG3 GTL2, FP504, prebp1, PRO0518, PRO2160, FLJ31163, FLJ42589  14q32  Imprinted Maternal
MEG8 Rian  14q32.31  Unknown Unknown
RTL1 MART1, PEG11, LOC388015  14q32.31 AS  Unknown Unknown
MAGEL2 nM15, NDNL1  15q11-q12 AS  Imprinted Paternal
NPAP1 C15orf2  15q11-q13  Imprinted Unknown
ZNF127AS MKRN3AS, Znp127as  15q11-q13  Unknown Unknown
MKRN3 D15S9, RNF63, ZFP127, ZNF127, MGC88288  15q11-q13  Imprinted Paternal
UBE3A AS, ANCR, E6-AP, HPVE6A, EPVE6AP, FLJ26981  15q11-q13 AS  Imprinted Maternal
SNRPN SMN, PWCR, SM-D, RT-LI, HCERN3, SNRNP-N, FLJ33569, FLJ36996, FLJ39265, MGC29886, SNURF-SNRPN, DKFZp762N022, DKFZp686C0927, DKFZp761I1912, DKFZp686M12165  15q11.2  Imprinted Paternal
SNORD116@ PET1, PWCR1, HBII-85  15q11.2  Imprinted Paternal
SNORD107 HBII-436, HBII-436 C/D box snoRNA  15q11.2  Imprinted Paternal
ATP10A ATPVA, ATPVC, ATP10C, KIAA0566  15q11.2 AS  Imprinted Maternal
SNORD109A HBII-438A  15q11.2  Imprinted Paternal
SNORD108 HBII-437, HBII-437 C/D box snoRNA  15q11.2  Imprinted Paternal
SNORD115-48 HBII-52-48  15q11.2  Imprinted Paternal
SNORD109B HBII-438B, HBII-438B C/D box snoRNA  15q11.2  Imprinted Paternal
PWCR1 PET1, non-coding RNA in the Prader-Willi critical region  15q11.2  Imprinted Paternal
SNORD115@ HBII-52  15q11.2  Imprinted Paternal
GABRA5 MGC138184  15q11.2-q12  Conflicting Data Paternal
GABRB3 MGC9051  15q11.2-q12 AS  Conflicting Data Paternal
NDN HsT16328  15q11.2-q12 AS  Imprinted Paternal
GABRG3   15q12  Conflicting Data Paternal
SNORD64 HBII-13, HBII-13 snoRNA  15q12  Imprinted Paternal
SNURF   15q12  Imprinted Paternal
GATM AT, AGAT  15q21.1 AS  Unknown Unknown
RASGRF1 GNRP, GRF1, CDC25, GRF55, CDC25L, H-GRF55, PP13187  15q24.2 AS  Unknown Unknown
MIR184 EDICT, MIRN184, miR-184  15q25.1  Unknown Unknown
SOX8 MGC24837  16p13.3  Predicted Paternal
ZNF597   16p13.3 AS  Imprinted Maternal
NAA60 HAT4, NAT15  16p13.3  Imprinted Maternal
SALL1 TBS, HSAL1, ZNF794  16q12.1 AS  Predicted Maternal
C16orf57 HVSL1, FLJ13154  16q21  Predicted Maternal
ACD PIP1, PTOP, TPP1, TINT1  16q22.1 AS  Predicted Maternal
FOXF1 FKHL5, ACDMPV, FREAC1, MGC105125  16q24  Predicted Maternal
TMEM88 FLJ20025, MGC71744  17p13.1  Predicted Maternal
PYY2   17q11  Predicted Paternal
HOXB2 K8, HOX2, HOX2H, Hox-2.8  17q21-q22 AS  Predicted Maternal
HOXB3 HOX2, HOX2G, Hox-2.7  17q21.3 AS  Predicted Maternal
LOC100131170 LOC100131170  17q25.3 AS  Predicted Paternal
IMPACT MGC33718  18q11.2-q12.1  Not Imprinted Biallelic
BRUNOL4 CELF4, BRUNOL-4  18q12 AS  Predicted Maternal
FAM59A GAREM, Gm944, C18orf11  18q12.1 AS  Predicted Paternal
TCEB3C HsT829, TCEB3L2, Elongin A3  18q21.1 AS  Imprinted Maternal
PPAP2C LPP2, PAP-2c, PAP2-g  19p13 AS  Predicted Maternal
DNMT1 AIM, DNMT, MCMT, CXXC9, HSN1E, ADCADN  19p13.2 AS  Imprinted Paternal
CHMP2A BC2, BC-2, VPS2, CHMP2, VPS2A  19q AS  Predicted Maternal
TSHZ3 TSH3, ZNF537, FLJ54422, KIAA1474  19q12 AS  Predicted Paternal
CHST8 GalNAc4ST, GALNAC4ST1  19q13.1  Predicted Maternal
ZNF225 MGC119735  19q13.2  Predicted Paternal
PEG3 PW1, ZNF904, ZSCAN24  19q13.4 AS  Imprinted Paternal
MIMT1 MIM1, LINC00067, NCRNA00067  19q13.4  Imprinted Paternal
ZIM2 ZNF656  19q13.4 AS  Imprinted Paternal
LILRB4 HM18, ILT3, LIR5, CD85K, LIR-5, LILRB5  19q13.4  Predicted Maternal
ZNF264 Zfp264  19q13.4  Unknown Unknown
MZF1 MZF-1, MZF1B, ZNF42, Zfp98, ZSCAN6  19q13.4 AS  Predicted Maternal
ZNF229 FLJ34222  19q13.31 AS  Predicted Maternal
NLRP2 NBS1, PAN1, NALP2, PYPAF2, CLR19.9  19q13.42  Imprinted Maternal
MIR371A C19MC, MIR371, MIRN371, hsa-mir-371, hsa-mir-371a  19q13.42  Imprinted Paternal
ZIM3 ZNF657  19q13.43 AS  Unknown Unknown
USP29 MGC163266, MGC163270, HOM-TES-84/86  19q13.43  Unknown Unknown
C20orf82 bA149I18.1, dJ1077I2.1  20p12.1  Predicted Paternal
ISM1 ISM, Isthmin, C20orf82, bA149I18.1, dJ1077I2.1  20p12.1  Predicted Paternal
PSIMCT-1   20q11.2  Imprinted Paternal
NNAT Peg5  20q11.2-q12  Imprinted Paternal
BLCAP BC10  20q11.2-q12 AS  Imprinted Isoform Dependent
MCTS2   20q11.21  Imprinted Paternal
HM13 H13, SPP, IMP1, PSL3, IMPAS, SPPL1, PSENL3, IMPAS-1, MSTP086, dJ324O17.1  20q11.21  Unknown Unknown
COL9A3 IDD, MED, EDM3, FLJ90759, DJ885L7.4.1  20q13.3  Predicted Maternal
GNAS AHO, GSA, GSP, POH, GPSA, NESP, GNAS1, PHP1A, PHP1B, C20orf45, MGC33735, dJ309F20.1.1, dJ806M20.3.3  20q13.3  Imprinted Isoform Dependent
L3MBTL L3MBTL1, FLJ41181, KIAA0681, H-L(3)MBT, dJ138B7.3, DKFZp586P1522  20q13.12  Imprinted Paternal
GNASAS SANG, NESPAS, GNAS1AS, NCRNA00075  20q13.32 AS  Imprinted Paternal
SANG SANG, Nespas  20q13.32  Imprinted Paternal
MIR296 MIRN296, miRNA296  20q13.32 AS  Imprinted Paternal
MIR298 MIRN298, hsa-mir-298  20q13.32 AS  Imprinted Paternal
C20orf20 Eaf7, MRGBP, URCC4, MRG15BP, FLJ10914  20q13.33  Predicted Maternal
SIM2 SIM, bHLHe15, MGC119447  21q22.2  Predicted Paternal
DGCR6L   22q11 AS  Imprinted Unknown
DGCR6   22q11.21  Imprinted Unknown
FLJ20464 FLJ20464  22q12.2  Predicted Paternal
TSIX XISTAS, NCRNA00013  Xq13.2  Not Imprinted Biallelic
XIST XCE, XIC, SXI1, swd66, DXS1089, DXS399E, NCRNA00001, DKFZp779P0129  Xq13.2 AS  Not Imprinted Biallelic